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There are five main types of agriculture that fungus-growing ants practice:[5] lower, coral fungi, yeast, generalized higher, and leafcutter agricultural systems. Lower agriculture is the most primitive system and is currently practiced by 80 species in 10 genera.[6][7] Coral-fungus agriculture is practiced by 34 species by a single derived clade within the genus Apterostigma.[7] The coral fungus farmers underwent a switch of cultivars between 10 and 20 million years ago to a non-leucocoprineacoeous fungus, which makes its choice of cultivar different from all other attines.[8][9] Yeast agriculture is practiced by 18 species of Cyphomyrmex rimosus. The C. rimosus group is hypothesized to have evolved growing fungus in a yeast form between 5 and 25 million years ago.[9] Generalized higher agriculture is practiced by 63 species in two genera and refers to the condition of highly domesticated fungus.[7] The fungi used in higher agriculture cannot survive without its agriculturalists to tend it and has phenotypic changes that allow for increased ease of ant harvesting.[9] Leafcutter agriculture, which is a more highly derived form of higher agriculture, is practiced by 40 species in two genera and has the most recent evolution, originating between 8 and 12 million years ago.[9] Leaf cutters use living biomass as the substrate to feed their fungi, whereas in all other types of agriculture, the fungus requires dead biomass.[9]