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[Spiderking50]

If i were to make the megabeast's region subterranean, would it only attack from the caverns? Or would it appear above ground when attacking? And would giving it a liar underground change that?

[i2amroy]

I believe it will still appear on the surface just like any other megabeast (and in fact you might have difficulties getting them to spawn at all).

Having lairs just determines what site type it lives in and has no real effect on how it attacks you.